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When Laying Tongue and Groove plywood over 1x6 subfloor, do I lay it with the grain perpendicular to the floor joists or to the 1x6 boards underneath?

I am redoing a bedroom and I am in the process of installing a new sub floor. I currently have 1x6 running perpendicular to the floor joists. Some do have a bit of give to them. I am laying over that ...